Skip to content

Free email delivery

Please sign up for email delivery in the subscription area to the right.
No salesman will call, at least not from us. Maybe from someone else.

Sandpipers & Other Goodies: Malibu Lagoon, 22 Sept. 2024

October 1, 2024

[Text by Chuck Almdale; photos by Elyse Jankowski, Lillian Johnson, Armando Martinez & Chris Tosdevin]

Don’t miss the quiz at the end which has some of the best photos.

White-faced Ibis (Elyse Jankowski 9-22-24)

The early arrivers to the lagoon got to see the ibis pair. I saw them fly away, but another one dropped in later. That’s migration – birds fly in, birds fly out, some stay and feed, some sleep. Our Greater Yellowlegs from last month was there, or more likely a new one. They show up only 4% of our visits but with two months in a row, they cranked it all the way up to a whopping 6.1%. Ibis are even rarer – just under 2%.

Morning snag-cormorants battled for perching room. (Lillian Johnson 9-22-24)

The lagoon outlet – open last month – had filled with sand. As a result the lagoon water level was quite high and didn’t fluctuate during our visit. One male Great-tailed Grackle strutted the sand. They’re 18″ long bill-tip to tail-tip, a half-inch longer than the American Crow, believe it or not. (It’s all in the tail.) The females are 3″ shorter; again, all in the tail. These birds nest somewhere in the general lagoon area, I’m not sure where. Perhaps up the creek, perhaps in the reed-bed across the lagoon by Adamson House.

Great-tailed Grackle (Chris Tosdevin 9-22-24)

Photographer Elyse Jankowski had to leave early but sent me some very nice photos. This Snowy Egret appears to have some Cattle Egret-like faint reddish highlights at the nape and lower back. I don’t know why, unless it was just a hint of mud.

Snowy Egret (Elyse Jankowski 9-22-24)

The Snowy Plovers have definitely returned! We had 42 birds, up from 22 in August and 6 in July. Two were banded, as photographed by Plover Wardens Larry Loeher and Grace Murayama three days earlier.

Western Snowy Plover, banded Pb:ob. You have to look carefully for the pink band. (Larry Loeher 9-19-24)
Western Snowy Plover, banded bg:rb. This juvenile was banded at hatch on 5/17/24 at Salinas River State Beach, and last seen around Monterey on July 22nd. (Larry Loeher 9-19-24)

We had some tricky times with several sandpipers, Least and Western (or Semipalmated).

The Western Below has a bill a bit shorter than usual, and close observation revealed semipalmated feet (check its right foot in 1st picture, left foot in 2nd picture). So – as its migration and just about anything can show up anywhere (within reason) – the possibility of an eastern Semipalmated Sandpiper reared its delicious head. We studied and scoped and discussed. We later studied the photos, even sending one or more to iNaturalist and waiting to see what the eBird people had to say. But, alas, it was not to be.

A Short Treatise on Sandpiper Toes

Birds have two toes (Ostrich), three toes (emus, bustards, the two species of Three-toed Woodpecker & quails), or four toes (everything else). Sandpipers have four toes arranged anisodactylly: three point forward, one points to the rear. They are numbered: #1 – back toe, #2 inner front, #3 middle front, #4 outer front. In photographs its often hard or impossible to see the #1 rear toe. Birds that spend their time in or near water often have webbing between their toes. Ducks, for example. Both the Semipalmated and Western Sandpipers have semipalmated feet – the webbing between their 4th and 3rd toes is larger than between the 3rd and 2nd toes.

Western Sandpiper, showing larger webbing between right foot 4th and 3rd toes. (Chris Tosdevin 9-22-24)

So the word “semipalmated” does describe the foot of Calidris pusilla, the Semipalmated Sandpiper, but it does not exclude Calidris mauri, the Western Sandpiper. And this can easily confuse the billions of people who expect descriptive animal names to a) exist, b) work, c) exclude, d) be useful, and e) not confuse. [A fool’s errand if ever there was one.] My understanding is that the Semipalmated Sandpipers were described first. When someone decided that the longer-billed and slightly-differently plumaged and shaped birds among them were actually a different species, the new birds were named “Western” as they were more numerous the farther west you went. [I wrote about this here in August 2022. Someday I hope to get all the facts, dates and name changes absolutely straight.] Just to confuse the issue, both species have variable bill lengths, with the Western a bit longer, but the westerly Semipalmated bill is a bit longer than the easterly Semipalmated bill.

Same Western Sandpiper, now showing larger webbing between left foot 4th and 3rd toes. (Chris Tosdevin 9-22-24)

But that was not the end of confusion and debate! The first bird disappeared, but we then thought we rediscovered it resting on the ground among the small pieces of driftwood.

Sandpiper X resting. (Chris Tosdevin 9-22-24)

Armando Martinez got a photo of it rising.

Sandpiper X rising. (Armando Martinez 9-22-24)

He then stretches his wings and staggers off. Not the back of the Snowy Plover and the head of the Black-bellied Plover behind him, providing some size comparison.

Sandpiper X has yellow-green legs. (Armando Martinez 9-22-24)
The Least Sandpiper makes his way down to the lagoon edge. (Chris Tosdevin 9-22-24)
Across the channel to Malibu Colony. A cormorant works to get off the water, pile-driver in the distance tries to root a not-yet-built house to the bedrock (is there any?). (Lillian Johnson 9-22-24)
Great Blue Heron (Elyse Jankowski 9-22-24)

Flowers are still blooming, or blooming again, in late September.

West Channel, back of Malibu Colony in distance. (Lillian Johnson 9-22-24)

This Pied-billed Grebe’s bill is not at all pied and it seems to have a wicked-looking thorny tip.

Pied-billed Grebe (Chris Tosdevin 9-22-24)

Last but not least, but without photo, we had a Lark Sparrow Chondestes grammacus in a very messy molting plumage, flitting around between the beach plants. I was very surprised to see it, as our only prior sighting of this species at the lagoon was on April 26, 1999, well back into the “before-times.”

As these photos are all taken this September at the lagoon, many of the birds are in plumage transition.

#1.
#2.
#3.
#4.
#5.
#6.
#7. Same individual. Could it be one of the Crombecs, flown over from West Africa?
#8.
#9. AKA Baldpate.
#10.
#11.
#12.
#13. At the lagoon 96% of the time.
#14. Count the legs, if you can.
#15.
#16. What are they, are the all the same species, how many?
#17. Is it a “sandpiper” or something else?
#18. Same question. And notice the sand.
#19.
#20.
#21.

Quiz Answers & credits
#1. Say’s Phoebe, first appearance since last November. (Chris Tosdevin 9-22-24)
#2. An exceptionally fierce-looking Wrentit (Elyse Jankowski 9-22-24)
#3. Blue-gray Gnatcatcher, increasingly common since 2015 (Chris Tosdevin 9-22-24)
#4. Savannah Sparrow, only the 4th sighting in the past 6 years. (Chris Tosdevin 9-22-24)
#5. Red-winged Blackbird: first year male, red not visible, brown bars on back. (Chris Tosdevin 9-22-24)
#6. Common Yellowthroat, male; breeds around the lagoon. (Chris Tosdevin 9-22-24)
#7. Great-tailed Grackle, male, whose tail has vanished (temporarily, we hope). (Grace Murayama 9-19-24)
#8. Ring-necked Ducks, 1 female, 3 males. Our 7th sighting at the lagoon. (Chris Tosdevin 9-22-24)
#9. American Wigeon, male, common winter visitor. (Chris Tosdevin 9-22-24)
#10. Pied-billed Grebe, a different view. (Larry Loeher 9-19-24)
#11. All-black Mallard. Not a male as no curly tail. If I’m wrong and this is not a mallard, let me know why. (Elyse Jankowski 9-22-24)
#12. Pygmy Blue Butterfly (Brephidium exilis), our lagoon specialty. (Chris Tosdevin 9-22-24)
#13. Great-blue Heron. (Chris Tosdevin 9-22-24)
#14. Whimbrel & 2 Willets. (Chris Tosdevin 9-22-24)
#15. Ruddy Turnstone, molting. (Chris Tosdevin 9-22-24)
#16. 27 Western Snowy Plovers (Chris Tosdevin 9-22-24)
#17. Semipalmated Plover (Chris Tosdevin 9-22-24)
#18. Black-bellied Plover (Elyse Jankowski 9-22-24)
#19. Greater Yellowlegs (Chris Tosdevin 9-22-24)
#20. Willet, molting. (Elyse Jankowski 9-22-24)
#21. Whimbrel (Elyse Jankowski 9-22-24)

Malibu Lagoon on eBird as of 9-27-24: 8053 lists, 2582 eBirders, 318 species.
Most recent species added: Cassin’s Vireo, 9/23/24 by Colin Drummond.

Many, many thanks to photographers: Elyse Jankowski, Lillian Johnson, Armando Martinez, Chris Tosdevin.

Upcoming SMBAS scheduled field trips; no reservations or Covid card necessary unless specifically mentioned:

  • Huntington Central Park, Sat. Oct 12, 8 am, contact leader Liz Galton (424-832-3504)
  • Malibu Lagoon, Sun. Oct 27, 8:30 (adults) & 10 am (parents & kids)
  • Franklin Canyon, Weds. Nov 13, 8:30 am. Our first mid-week walk, enter at south end.
  • These and any other trips we announce for the foreseeable future will depend upon expected status of the Covid/flu/etc. pandemic at trip time. Any trip announced may be canceled shortly before trip date if it seems necessary. By now any other comments should be superfluous.
  • Link to Programs & Field Trip schedule.

The next SMBAS Zoom program: Professor Barney Schlinger, UCLA Dept. of Ecology & Evolutionary Biology; Manakins, Evening Meeting, Tuesday, October 8, 2024, 7:30 p.m.

The SMBAS 10 a.m. Parent’s & Kids Birdwalk is again running. Reservations not necessary for families, but for groups (scouts, etc.), call Jean (213-522-0062).

Links: Unusual birds at Malibu Lagoon
9/23/02 Aerial photo of Malibu Lagoon
More recent aerial photo

Prior checklists:
2023: Jan-June, July-Dec 2024: Jan-June
2021: Jan-JulyJuly-Dec2022: Jan-June, July-Dec
2020: Jan-JulyJuly-Dec  2019: Jan-June, July-Dec  
2018: Jan-June, July-Dec  2017: Jan-June, July-Dec
2016: Jan-June, July-Dec  2015: Jan-May, July-Dec
2014: Jan-July,  July-Dec  2013: Jan-June, July-Dec
2012: Jan-June, July-Dec 2011: Jan-June, July-Dec
2010: Jan-June, July-Dec  2009: Jan-June, July-Dec

The 10-year comparison summaries created during the Lagoon Reconfiguration Project period, remain available—despite numerous complaints—on our Lagoon Project Bird Census Page. Very briefly summarized, the results unexpectedly indicate that avian species diversification and numbers improved slightly during the restoration period June’12-June’14.

Many thanks to Marie Barnidge-McIntyre, Femi Faminu, Elyse Jankowski, Lillian Johnson, Ruth & Chris Tosdevin & others for their contributions to this month’s checklist.

The species lists below is irregularly re-sequenced to agree with the California Bird Records Committee Official California Checklist. If part of the right side of the chart below is hidden, there’s a slider button inconveniently located at the bottom end of the list. The numbers 1-9 left of the species names are keyed to the nine categories of birds at the bottom.
[Chuck Almdale]

Malibu Census 20244/285/266/237/288/259/22
Temperature62-7257-6462-7263-7264-7867-75
Tide Lo/Hi HeightL-0.14L-0.77L-1.17L+1.81L+2.00L+2.09
 Tide Time073806350537091607350611
1Brant (Black) 2    
1Canada Goose6894  
1Northern Shoveler 2    
1Gadwall202235271230
1American Wigeon     1
1Mallard101415628
1Ring-necked Duck     4
1Surf Scoter4     
1Long-tailed Duck1     
1Red-breasted Merganser42    
2Pied-billed Grebe 11115
2Western Grebe 91   
7Feral Pigeon 53691
7Mourning Dove 25231
8Anna’s Hummingbird221 2 
8Allen’s Hummingbird151234
2American Coot 1  272
5Black-necked Stilt  2   
5Black Oystercatcher4     
5Black-bellied Plover  1517067
5Killdeer64312412
5Semipalmated Plover9   133
5Snowy Plover   62242
5Whimbrel42 5222
5Long-billed Curlew   11 
5Wilson’s Phalarope  1   
5Spotted Sandpiper1  2 1
5Willet2 2955
5Greater Yellowlegs1   11
5Ruddy Turnstone    13
5Sanderling    21
5Least Sandpiper2   233
5Western Sandpiper   6132
6Bonaparte’s Gull210    
6Heermann’s Gull 66542524
6Ring-billed Gull43  22
6Western Gull164516022011355
6California Gull60383102375
6Glaucous-winged Gull   11 
6Caspian Tern208 1441
6Forster’s Tern  1 2 
6Elegant Tern2001902524010 
6Royal Tern602  55
2Red-throated Loon2     
2Common Loon 1    
2Brandt’s Cormorant 35    
2Pelagic Cormorant 14 1 
2Double-crested Cormorant2612024283754
2Brown Pelican2353481251632724
3Black-crowned Night-Heron112111
3Snowy Egret327434
3Green Heron  1112
3Great Egret549332
3Great Blue Heron4 3342
3White-faced Ibis     3
4Turkey Vulture 1 417
4Osprey1     
4Red-shouldered Hawk     1
4Red-tailed Hawk     2
8Belted Kingfisher    11
8Nuttall’s Woodpecker 1   1
8Hairy Woodpecker    1 
8Nanday Parakeet     25
9Black Phoebe473632
9Say’s Phoebe     1
9Ash-throated  Flycatcher    1 
9Cassin’s Kingbird4   1 
9Western Kingbird    8 
9Warbling Vireo 1    
9California Scrub-Jay1    2
9American Crow346448
9Common Raven23    
9Oak Titmouse    1 
9Tree Swallow  3  1
9Violet-green Swallow28    
9No. Rough-winged Swallow525 2 
9Barn Swallow10202020201
9Cliff Swallow 306 4 
9Bushtit25919520
9Wrentit 21  4
9Ruby-crowned Kinglet     1
9Blue-gray Gnatcatcher    14
9House Wren 1  23
9Marsh Wren     2
9Bewick’s Wren     2
9European Starling    110
9Northern Mockingbird    1 
9Scaly-breasted Munia  1   
9House Finch1015111254
9Lesser Goldfinch252  3
9Lark Sparrow     1
9Dark-eyed Junco 2  12
9Savannah Sparrow     1
9Song Sparrow105541010
9California Towhee14411 
9Spotted Towhee  1   
9Western Meadowlark     15
9Hooded Oriole  111 
9Red-winged Blackbird  11  11
9Brown-headed Cowbird    18 
9Great-tailed Grackle 233 1
9Orange-crowned Warbler11  12
9Common Yellowthroat 1  15
9Yellow Warbler     1
9Townsend’s Warbler     2
Totals by TypeAprMayJunJulAugSep
1Waterfowl455059371443
2Water Birds – Other26351615519268155
3Herons, Egrets & Ibis13722121214
4Quail & Raptors1104110
5Shorebirds2969139157142
6Gulls & Terns362302254527165162
7Doves0788122
8Other Non-Passerines3822731
9Passerines57118927092119
 Totals Birds7731015601991528678
        
 Total SpeciesAprMayJunJulAugSep
1Waterfowl663324
2Water Birds – Other385354
3Herons, Egrets & Ibis435556
4Quail & Raptors110113
5Shorebirds82581212
6Gulls & Terns785696
7Doves022222
8Other Non-Passerines232144
9Passerines14191792227
Totals Species455244386268

Coastal Cleanup Day Reminder – Sat. Sept. 21 – 9am to noon – Malibu Lagoon

September 20, 2024

Saturday, Sept. 21, 2024: Annual International Coastal Cleanup Day, from 9:00 A.M. to Noon at the Malibu Lagoon.  Help us clear the trash around the lagoon!  Last year, more than 7,000 volunteers in the Los Angeles area removed over 16,000 lbs. of trash and 400 lbs. of recycling from 97 miles of beach, river, underwater, and trail cleanup sites! Ninety percent of all floating marine debris is plastic.  As we know, bright colored plastics or small micro-plastics can be confused for food.  A 2012 study by the Convention on Biological Diversity found that 663 marine species have been impacted by plastic litter through ingestion or entanglement.  It is important that we clean the lagoon area before the first rains come and carry everything out to the ocean.

Chris deals with weighty matters (L.Johnson 9/20/14)

Chris deals with weighty matters (L.Johnson 9/20/14)

Registration begins at 8:30 a.m.  We encourage you to get waivers and registration forms on-line at https://healthebay.org/coastalcleanupday/. Scroll down to “Register Now” button and click it, then scroll down to “Join a Cleanup Site” button and click it, then scroll down to Malibu Lagoon State Beach, click the number of people coming, then click the “Join a Cleanup” button, then fill out the registration form, then click the “Next” or “Save” button and finish the process.

Liability waivers (different form from registration) will be available at the site for you to sign. 

Our chapter concentrates its efforts at Malibu Lagoon, but you can call 1-800-HEALBAY for information and other places to volunteer.  Parking passes will available at the lagoon on this day.  If possible, bring your own gloves, bucket for trash,  and sunscreen. Don’t worry if you forget such stuff because from 9:00 a.m. until noon, volunteers will be given supplies and instructions on how to carry out a beach cleanup.

Family Guide: Suitable for everyone but toddlers.  Small children, already built conveniently close to the ground, are great at picking up those tiny pieces of plastic.

Information Contact: Jean Garrett (213-522-0062)

[Directions] Malibu Lagoon is at the intersection of Pacific Coast Highway and Cross Creek Road in Malibu.  Parking in the official lagoon lot is normally $3/hour (should be free today) or by annual pass.  You may also park either along PCH north of Cross Creek Road or on Cross Creek Road itself but be careful – some parts of PCH are off-limits (read the signs carefully.)  Lagoon parking in the shopping center lot is not permitted.

Hey! Look guys! You can see the bottom! (J Kenney)

Hey! Look guys! You can see the bottom! (J Kenney)

Avian Checklist of 41 Orders & 251 Families | Taxonomy 10

September 12, 2024

[By Chuck Almdale]

Introduction

My research could uncover no single source for the most up-to-date taxonomic information on birds (or anything else); even if they were such a thing, there is such wide disagreement among those working on the avian phylogenetic sequence that it could be biased and a source more for dissension than agreement. One such source may exist that everyone in the world agrees is completely accurate and completely up-to-date, but if there is, it’s a very well-kept secret.

Therefore, the following taxonomic sequence draws from multiple sources, most notably Birds of the World (BOW) website – Orders & Families, Distribution and Taxonomy of Birds of the World (Sibley & Monroe, 1990), several papers especially “The Origin and Diversification of Birds” (Brusatte, O’Conner & Jarvis, 2014) [Link to Article] and numerous Wikipedia pages, especially Passerine and Passerea  and the separate pages for their nearly innumerable subdivisions. [Wikipedia is remarkably unbiased, and their articles frequently offer multiple taxonomic sequences where disagreement exists.] Research continues; refinements and rearrangements will continue to appear as research is completed; gradually a unified picture is developing. The following sequence is as accurate as I could make it at this time. What follows are some guidelines as to what is in the sequence and a few choices I made for the sake of presentation.

All 41 orders and 251 families are numbered according to the BOW Orders & Families as of mid-June 2024, which may well have changed by the time this gets posted. I expect some major changes to this sequence to occur within the coming year as the International Ornithological Union (IOU) Working Group Avian Checklist Committees [Link] complete their work, or at least post some updates. Following this, eBird/Birds of the World will undoubtedly update their sequences, as they regularly do at least yearly as the Clements List is updated. It may come to pass within the next few years that the Clements and IOU checklists are brought more into alignment, particularly in regards to the higher levels of taxonomy (from Class down to Family).

When Sibley & Monroe published their book in 1990, their table of contents utilized many additional taxonomic ranks that possessed useful suffixes (in brackets below) for uniformity in their nineteen taxonomic ranks:  
Class: Super-, Class, Sub-, Infra- (-es), Parv- (-ae)
Order: Super- (-imorphae), Order (-iformes), Sub- (-i), Infra- (-ides), Parv- (-ida)
Family: Super- (-oidea), Family (-idae), Sub- (-inae), Tribe (-ini), Subtribe (-ina)
Genus: Genus, subgenus
Species: species, subspecies

These ranks are still used in many books, papers and websites and I retained them whenever possible. But a lot of taxonomic research, particularly in the development of evolutionary clocks, has occurred in the 34 years since that book was published.

Clades

The use of clades and clade terminology has grown enormously over the past few decades. Many of the previous terms have been adopted into the clade structure, especially Order, Family, Genus and Species, but also many of the subdivisions listed above. The clade structure, which works far better in cladogram diagrams than in writing, emerged out of the growing body of information concerning divergences in evolutionary lineages and the development and availability of “evolutionary clocks.” When an evolutionary line divides, a “node” with two branches is created at that point which can be dated by means of the evolutionary clock. The terms Order, Suborder, etc. worked well for creating a hierarchy of ranks, but the number of nodes of division and clade ranks have already vastly outrun this restricted terminology, and the number of ranks can only continue to grow. Every time a species evolves into two subspecies than evolve farther apart into two species, a node is created and should be given a name. From the level of Class Aves down to Passer domesticus domesticus, there are at least seventeen named Linnaean-style ranks. Now, as we saw in Taxonomy 6, Clade63 Subclass Neornithes is only the beginning of extant modern birds. The cladogram in Taxonomy 9 added another 10 ranks to get to Order Passeriformes, and Sibley & Monroe add eight more ranks to arrive at Passer domesticus domesticus. That’s a total of 81 ranks, so far. Once the sequence to P.d.domesticus is fully mapped – and ranks are continually inserted all along the sequence – there could be well over a hundred ranks. I sure Linnaeus, who thought 10,000 species would cover the lot of living things, never conceived of such complexity even in his most fevered dreams of discovery and classification. People are already working on developing taxonomic modifications or replacement systems.

Although there are still nodes where there are more than two branches (daughter lineages), it is widely viewed that such polytomies are an artifact of insufficient information. When an evolutionary lineage branches, the process involves a single species branching into two species only, not three or more. As researchers produce finer-grained results, it is expected that all such polytomies will be replaced with a series of paired branches.

A good system of upper-level nomenclature could utilize a combination of names and numbers. If we kept only Class, Order, Family, Genus, Species we could use have Class 1, class 2, etc.; Order 1, order 2, etc.; Family 1, family 2, etc.;…genus 1, genus 2, etc. Class 1 = the original level of class, Class 2 = the next step down…Class 10 = the 9th level below Class 1, and so on. They could adopt subscripts as I did in prior postings, e.g. Class1, Class2…Class63, although, frankly, typing out all those subscripts was a nuisance. There would be no limit to the number of steps which could be added as necessary when new divergences are discovered. Currently many new clades are given names, but many aren’t, and you can find many cladograms where nodes and their daughter lineages are not named. The previous names from Infraclass to Subtribe had their unique suffixes, which was a good clue as to where they fell in the sequence (Passer-imorphae, Passer-iformes, Passer-idae, Passer) but most of the new clade names, if they are even named, give no linguistic clue as to their position in the sequence. One can see the nodes on a cladogram, but not in the written descriptions. A simple – yet sufficiently descriptive – system needs to be developed. The portmanteau clade names utilizing the next two taxa to form the new taxon name is a good start (e.g. Xenacoelomorpha: Xena + coel + omorpha, combines the two daughter ranks xenoturbella and acoelomorpha).

Key to the Avian Taxonomic List Below

Totals of Orders, Families and Species
1. Most ranks have a set of numbers indicating the number of major subdivisions (Order, Family, Species).
Subclass: Neornithes 41/251/11017 means it has 41 orders, 251 families, 11,017 species.  
2. Orders have two numbers for families and species:
18. Order: Gruiformes – Cranes & allies – 6/192 means that Order 18 Gruiformes has 6 families, 192 species.
3. Families show a single number for species: 123. Family:
Viduidae – Indigobirds – 20
means that family 123 Viduidae has 20 species.
4. Most intermediate steps are similarly numbered, although a few unnamed clades are not.
5. Many ranks have used (or are still using) multiple names (e.g. Superorder Caprimulgimorphae = Clade Strisores). I did not search for these alternate names, but where I encountered them, I included them in parentheses).

Bold, Italics & Indentation
Bold
and italics to differentiate higher-level and lesser-known sequence steps from the more common ranks. Each lower taxonomic rank is indented 3 spaces. Equal indentation means equivalent rank (two orders, two families, etc.). Indentation is recalibrated twice (locations noted within the sequence) to avoid word-wrapping.

Colors
The forty-one orders from the Chart of Five Checklists and the prior posting’s cladogram are numbered and highlighted in blue.
The nine major clades from the prior posting’s cladogram are highlighted in red.
I included a few families (unnumbered and highlighted in blue) that I think are likely to be added within the coming year, but did not include the probable number of the species classified to them as they are counted within their current families.

Family Number
Each family (e.g. 1. Family: Rheidae – Rheas) is given the sequence number as shown on the BOW website – Orders & Families. Sometimes, primarily in Passeriformes, their sequence in this list within (and even between) groups is not in numerical order. This occurs where I have strong reason to believe that sequences other than the BOW are more up-to-date and the BOW sequence will likely soon change.

The top levels (Domain to Class) were discussed in the earlier postings in this series.

You may find it interesting to compare the following written sequence to the cladogram included in Taxonomy 9. They agree to one other, but each includes information not contained within the other. Working together, they demonstrate the different strengths each form of presentation possesses.

If you’d like an 8-page PDF document of the entire sequence, click below.
Taxonomic Sequence of Avian 41 Orders and 251 Families

Family Size

It may or may not signify anything to note that most of the 251 bird families contain few species, only a few contain many dozens or hundreds of species. I noticed this when I discovered that 37 families (15.5% of 251 families) contained only a single species. As this was roughly 1/6th of the total, I divided the families into six groups based on species totals. One sixth of families are monotypic, one-sixth have more than 80 species each. It’s interesting to speculate on this peculiar fact as you go through the entire family list. What is it in the evolutionary process that makes so many species sufficiently different from all other species to warrant their own family? Do large families signify relative evolutionary success, adaptability or greater speed of speciation? Are monotypic families evolutionarily closer to the root of their clade?

The first extant species in the list.


The heaviest, tallest and probably most ancient bird in the world, the Common Ostrich, Struthio camelus. Photo: Yathin S Krishnappa. Wikipedia: Ostrich

The Taxonomic Sequence of Avian Orders and Families

To conserve horizontal space, Parvclass Neoaves is moving 21 spaces to the left.

Parvclass Neoaves – 34/238/10475
   Clade Columbea – 5/5/400
                  Clade Superorder: Phoenicopterimorphae – 2/2/28 (or Clade Mirandornithes)
                           8. Order: Phoenicopteriformes – 1/6
                                 1. Family: Phoenicopteridae – Flamingos – 6
                           9. Order: Podicipediformes – 1/22
                                 1. Family: Podicipedidae – Grebes – 22
                  Clade Superorder: Columbimorphae – 3/3/372
                           10. Order: Columbiformes – 1/353
                                 1. Family: Columbidae – Pigeons & Doves – 353
                           11. Order: Mesitornithiformes – 1/3
                                 1. Family: Mesitornithidae – Mesites – 3
                           12. Order: Pterocliformes – 1/16
                                 1. Family: Pteroclidae – Sandgrouse – 16
   Clade Passerea – 29/233/10075
      Clade Otidae – 4/11/805
                  Clade Superorder: Otidimorphae – 3/3/196
                           13. Order: Otidiformes – 1/26
                                 1. Family: Otididae – Bustards – 26
                           14. Order: Musophagiformes – 1/23
                                 1. Family: Musophagidae – Turacos – 23
                           15. Order: Cuculiformes – 1/147
                                 1. Family: Cuculidae – Cuckoos – 147
                  Superorder: Caprimulgimorphae – 1/8/609 (Clade Strisores)
                           16. Order: Caprimulgiformes – Nightbirds, Swifts, Hummers – 8/609
                                 2. Family: Caprimulgidae – Nightjars & Allies – 97
                        Clade Vanescaves
                                 3. Family: Nyctibiidae – Potoos – 7
                                 4. Family: Steatornithidae – Oilbird – 1
                        Clade Letornithes
                                 1. Family: Podargidae – Frogmouths – 16
                        Clade Daedalornithes
                                 5. Family: Aegothelidae – Owlet-nightjars – 9
                           Clade Apodiformes
                              Clade Suborder Apodi
                                 6. Family: Apodidae – Swifts – 112

                                 7. Family: Hemiprocnidae – Treeswifts – 4
                              Clade Suborder Trochili
                                 8. Family: Trochilidae – Hummingbirds – 363
      Clade Gruae – 3/26/579
                  Clade Superorder: Opisthocomimorphae – 1/1/1
                           17. Order: Opisthocomiformes – 1/1

                                 1. Family: Opisthocomidae – Hoatzin – 1
                  Clade Superorder: Gruimorphae 2/25/578 (or Charadriimorphae, Cursorimorphae)
                           18. Order: Gruiformes – Cranes & allies – 6/192
                              Suborder: Ralli – 3/173
                                 1. Family: Sarothruridae – Flufftails – 15
                                 2. Family: Rallidae – Rails, Gallinules & Coots – 155
                                 3. Family: Heliornithidae – Finfoots – 3
                              Suborder: Grui – 3/19
                                 4. Family: Aramidae – Limpkin – 1
                                 5. Family: Psophiidae – Trumpeters – 3
                                 6. Family: Gruidae – Cranes – 15
                           19. Order: Charadriiformes – Shorebirds & Gulls – 19/386
                              Suborder: Charadrii – 8/105
                                 1. Family: Pluvianellidae – Magellanic Plover – 1
                                 2. Family: Chionidae – Sheathbills – 2
                                 3. Family: Burhinidae – Thick-knees – 10
                                 4. Family: Pluvianidae – Egyptian Plover – 1
                                 5. Family: Recurvirostridae – Stilts & Avocets – 9
                                 6. Family: Ibidorhynchidae – Ibisbill – 1
                                 7. Family: Haematopodidae – Oystercatchers – 12
                                 8. Family: Charadriidae – Plovers & Lapwings – 69
                              Suborder: Scolopaci – 5/113
                                 9. Family: Pedionomidae – Plains-wanderer – 1
                                 10. Family: Thinocoridae – Seedsnipes – 4
                                 11. Family: Rostratulidae – Painted-Snipes – 3
                                 12. Family: Jacanidae – Jacanas – 8
                                 13. Family: Scolopacidae – Sandpipers & Allies – 97
                              Suborder: Lari – 6/168
                                 14. Family: Turnicidae – Buttonquail – 18
                                 15. Family: Dromadidae – Crab-Plover – 1
                                 16. Family: Glareolidae – Pratincoles & Coursers – 17
                                 17. Family: Stercorariidae – Skuas & Jaegers – 7
                                 18. Family: Alcidae – Auks, Murres & Puffins – 25
                                 19. Family: Laridae – Gulls, Terns & Skimmers – 100
      Clade Superorder Eurypygimorphae – 2/3/5 (or Phaethontimorphae)
                           20. Order: Eurypygiformes – 2/2
                                 1. Family: Rhynochetidae – Kagu – 1
                                 2. Family: Eurypygidae – Sunbittern – 1
                           21. Order: Phaethontiformes – 1/3
                                 1. Family: Phaethontidae – Tropicbirds – 3
      Clade Aequornithes – Core Waterbirds – 6/16/365
                  Clade Superorder: Gaviimorphae – 1/1/5  
                           22. Order: Gaviiformes – 1/5

                                 1. Family: Gaviidae – Loons – 5
         Clade Feraequornithes – 5/15/360
                  Clade Austrodyptornithes – 2/5/164 (or Superorder Procellariimorphae)

                           23. Order: Sphenisciformes – 1/18
                                 1. Family: Spheniscidae – Penguins – 18
                           24. Order: Procellariiformes – 4/146
                                 1. Family: Diomedeidae – Albatrosses – 20
                                 2. Family: Oceanitidae – Southern Storm-Petrels – 10
                                 3. Family: Hydrobatidae – Northern Storm-Petrels – 18
                                 4. Family: Procellariidae – Shearwaters & Petrels – 98
                  Clade Superorder: Pelecanimorphae – 3/10/196
                           25. Order: Ciconiiformes – 1/20

                                 1. Family: Ciconiidae – Storks – 20
                  Clade: Pelecanes – 2/9/176
                           26. Order: Suliformes – 4/59
                                 1. Family: Fregatidae – Frigatebirds – 5
                                 2. Family: Sulidae – Boobies & Gannets – 10
                                 3. Family: Anhingidae – Anhingas – 4
                                 4. Family: Phalacrocoracidae – Cormorants & Shags – 40
                           27. Order: Pelecaniformes – 5/117
                              Suborder: Pelicani – 3/10
                                 1. Family: Pelecanidae – Pelicans – 8
                                 2. Family: Balaenicipitidae – Shoebill – 1
                                 3. Family: Scopidae – Hamerkop – 1
                              Suborder: Ardei – 2/107 
                                 4. Family: Ardeidae – Herons, Egrets & Bitterns – 71
                                 5. Family: Threskiornithidae – Ibises & Spoonbills – 36
      Clade Telluraves (Core Landbirds) – 14/177/8321  
         Clade Infraclass Afroaves – 10/28/1254
            Clade Superorder: Accipitrimorphae – 2/4/259
                           28. Order: Cathartiformes – 1/7

                                 1. Family: Cathartidae – New World Vultures – 7
                           29. Order: Accipitriformes – 3/252
                                 1. Family: Sagittariidae – Secretarybird – 1
                                 2. Family: Pandionidae – Osprey – 1
                                 3. Family: Accipitridae – Hawks, Eagles & Kites – 250
            Clade Superorder: Strigimorphae – 1/2/247
                           30. Order: Strigiformes – 2/247

                                 1. Family: Tytonidae – Barn-Owls – 18
                                 2. Family: Strigidae – Owls – 229
            Clade Superorder: Coriciimorphae – 7/22/748
                           31. Order: Coliiformes – 1/6

                                 1. Family: Coliidae – Mousebirds – 6
               Clade Cavitaves – 6/21/742
                           32. Order: Leptosomiformes – 1/1

                                 1. Family: Leptosomidae – Cuckoo-roller – 1
                  Clade Eucavitaves – 5/20/741
                           33. Order: Trogoniformes – 1/46

                                 1. Family: Trogonidae – Trogons – 46
                     Clade Picocoraciae – 4/19/695
                           34. Order: Bucerotiformes – 4/75

                              Suborder: Upupi – 2/11
                                 1. Family: Upupidae – Hoopoes – 3
                                 2. Family: Phoeniculidae – Woodhoopoes & Scimitarbills – 8
                              Suborder: Buceroti – 2/64
                                 3. Family: Bucorvidae – Ground-Hornbills – 2
                                 4. Family: Bucerotidae – Hornbills – 62
                        Clade Picodynastornithes – 3/15/620 (was Superorder: Coraciimorphae)
                           35. Order: Coraciiformes – 6/185
                              Clade unnamed
                                 1. Family: Todidae – Todies – 5
                                 2. Family: Momotidae – Motmots – 14
                                 3. Family: Alcedinidae – Kingfishers – 117
                              Clade unnamed
                                 4. Family: Meropidae – Bee-eaters – 31
                                 5. Family: Coraciidae – Rollers – 13
                                 6. Family: Brachypteraciidae – Ground-Rollers – 5
                           36. Order: Galbuliformes (sometimes suborder Galbuli in Picaformes) – 2/55
                                 1. Family: Bucconidae – Puffbirds – 37
                                 2. Family: Galbulidae – Jacamars – 18
                           37. Order: Piciformes – 7/380
                              Infraorder Ramphastides – 5/129
                                 1. Family: Lybiidae – African Barbets – 41
                                 2. Family: Megalaimidae – Asian Barbets – 35
                                 3. Family: Capitonidae – New World Barbets – 15
                                 4. Family: Semnornithidae – Toucan-Barbets – 2
                                 5. Family: Ramphastidae – Toucans – 36
                              Infraorder: Picides – 2/251
                                 6. Family: Indicatoridae – Honeyguides – 16
                                 7. Family: Picidae – Woodpeckers – 235
         Clade Infraclass Australaves – 4/149/7067
                           38. Order: Cariamiformes – 1/2
                                 1. Family: Cariamidae – Seriemas – 2
            Clade Superorder Eufalconimorphae – 148/7065   
                           39. Order: Falconiformes – 1/65

                                 1. Family: Falconidae – Falcons & Caracaras – 65
                  Clade Psittacopasseres – 147/7000 (formerly Passerimorphae)
                           40. Order: Psittaciformes – 4/405
                              Superfamily: Strigopoidea – 1/4
                                 1. Family: Strigopidae – New Zealand Parrots – 4
                              Superfamily: Cacatuoidea – 1/22
                                 2. Family: Cacatuidae – Cockatoos – 22
                              Superfamily: Psittacoidea – True Parrots – 2/379
                                 3. Family: Psittaculidae – Old World Parrots – 202
                                 4. Family: Psittacidae – New World & African Parrots – 177
                           41. Order: Passeriformes – 143/6595

The final extant species in the non-passerine list.


Hispaniolan Parakeet, Psittacara chloropterus.
Photo: Martingloor. Wikipedia: Hispaniolan Parakeet

To conserve horizontal space, Order Passeriformes is moving 27 spaces to the left. 

The first extant species in the passerine list.


 Rifleman, female, Acanthisitta chloris, of South Island, New Zealand. An insectivorous wren-like member of a four-species family basal to all passerines, endemic to New Zealand. Photo: digitaltrails. Wikipedia: Rifleman 

As you go through the following list, note the 3 suborders, 4 infraorders, 5 parvorders and 10 superfamilies.

41. Order: Passeriformes – 143/6595
   Suborder Acanthisitti – 1/4
               1. Family: Acanthisittidae – New Zealand Wrens – 4
   Suborder
Tyranni – Non-oscine Passerines – 17/1386
     
Infraorder Eurylaimides – Old World Suboscines – 5/68

               5. Family: Philepittidae – Asities – 4
               3. Family: Eurylaimidae – Asian & Grauer’s Broadbills – 10
               2. Family: Calyptomenidae – African & Green Broadbills – 6
               4. Family: Sapayoidae – Sapayoa – 1
               6. Family: Pittidae – Pittas – 47
     
Infraorder Tyrannides – New World Suboscines – 12/1318

         Parvorder Furnariida – 7/714
               8. Family: Melanopareiidae – Crescentchests – 4
               9. Family: Conopophagidae – Gnateaters – 12
               7. Family: Thamnophilidae – Typical Antbirds – 237
               10. Family: Grallariidae – Antpittas – 70
               11. Family: Rhinocryptidae – Tapaculos – 65
               12. Family: Formicariidae – Antthrushes – 12
               13. Family: Furnariidae – Ovenbirds & Woodcreepers – 314
         Parvorder Tyrannida  5/604
               14. Family: Pipridae – Manakins – 55

               15. Family: Cotingidae – Cotingas – 65
               16. Family: Tityridae – Tityras and Allies – 35
               17. Family: Oxyruncidae – Sharpbill, Royal Flycatcher & Allies – 8
                     Family: Onychorhynchidae–Royal Flycatcher–Part of Oxyruncidae
               18. Family: Tyrannidae – Tyrant Flycatchers – 441
   Suborder Passeri – Oscine Passerines  125/5205
      Basal families in Passeri clade – 11/342
               20. Family: Atrichornithidae – Scrub-birds – 2
               19. Family: Menuridae – Lyrebirds – 2
               22. Family: Climacteridae – Australasian Treecreepers – 7
               21. Family: Ptilonorhynchidae – Bowerbirds – 27
               23. Family: Maluridae – Fairywrens – 32
               25. Family: Dasyornithidae – Bristlebirds – 3
               26. Family: Pardalotidae – Pardalotes – 4
               27. Family: Acanthizidae – Thornbills & Allies – 66
               24. Family: Meliphagidae – Honeyeaters – 191
               28. Family: Pomatostomidae – Pseudo-Babblers – 5
               29. Family: Orthonychidae – Logrunners – 3
      Infraorder Corvides30/847 (previously parvorder Corvida)
        Basal families in Corvides clade – 4/107
               30. Family: Cinclosomatidae – Quail-thrushes & Jewel-babblers – 12
               31. Family: Campephagidae – Cuckooshrikes – 89 (some in Orioloidea)
               32. Family: Mohouidae – Whiteheads – 3
               33. Family: Neosittidae – Sittellas – 3
        Superfamily: Orioloidea  8/180 
               34. Family: Psophodidae – Whipbirds & Wedgebills – 5
               35. Family: Eulacestomatidae – Ploughbill – 1
               37. Family: Falcunculidae – Shrike-tits – 3
               36. Family: Oreoicidae – Australo-Papuan Bellbirds – 3
               38. Family: Paramythiidae – Tit Berrypecker & Crested Berrypecker – 3
               39. Family: Vireonidae – Vireos, Shrike-Babblers & Erpornis – 61
               40. Family: Pachycephalidae – Whistlers & Allies – 63
               41. Family: Oriolidae – Old World Orioles – 41
            Superfamily: Malaconotoidea – 8/154
               42. Family: Machaerirhynchidae – Boatbills – 2
               43. Family: Artamidae – Woodswallows, Bellmagpies & Allies – 24
               44. Family: Rhagologidae – Mottled Berryhunter – 1
               49. Family: Malaconotidae – Bushshrikes & Allies – 50
               47. Family: Pityriasidae – Bristlehead – 1
               48. Family: Aegithinidae – Ioras – 4
               45. Family: Platysteiridae – Wattle-eyes & Batises – 32
               46. Family: Vangidae – Vangas, Helmetshrikes & Allies – 40
            Superfamily: Corvoidea – 10/406
               50. Family: Rhipiduridae – Fantails – 64
               51. Family: Dicruridae – Drongos – 28
               54. Family: Monarchidae – Monarch Flycatchers – 100
               53. Family: Ifritidae – Ifrita – 1
               52. Family: Paradisaeidae – Birds-of-Paradise – 44
               55. Family: Corcoracidae – White-winged Chough & Apostlebird – 2
               56. Family: Melampittidae – Melampittas – 2
               58. Family: Laniidae – Shrikes – 34
               57. Family: Platylophidae – Crested Shrikejay – 1
               59. Family: Corvidae – Crows, Jays & Magpies – 130
      Infraorder Passerides – 84/4016 (previously parvorder Passerida) 
         Basal families in Passerides clade – 8/76
               60. Family: Cnemophilidae – Satinbirds – 3
               61. Family: Melanocharitidae – Berrypeckers & Longbills – 12
               62. Family: Callaeidae – Wattlebirds – 5
               63. Family: Notiomystidae – Stitchbird – 1
               64. Family: Petroicidae – Australasian Robins – 50
               67. Family: Eupetidae – Rail-babbler – 1
               65. Family: Picathartidae – Rockfowl – 2
               66. Family: Chaetopidae – Rockjumpers – 2
         Parvorder Sylviida – 26/1363
            Basal families in Sylviida clade – 9/364
               68. Family: Hyliotidae – Hyliotas – 4
               69. Family: Stenostiridae – Fairy Flycatchers – 9
               70. Family: Paridae – Tits, Chickadees & Titmice – 63
               71. Family: Remizidae – Penduline-Tits – 11
               73. Family: Panuridae – Bearded Reedling – 1
               72. Family: Alaudidae – Larks – 93
               74. Family: Nicatoridae – Nicators – 3
               75. Family: Macrosphenidae – African Warblers – 18
               76. Family: Cisticolidae – Cisticolas & Allies – 162
            Superfamily: Locustelloidea – 6/231
               77. Family: Acrocephalidae – Reed Warblers & Allies – 60
               78. Family: Locustellidae – Grassbirds & Allies – 67
               79. Family: Donacobiidae – Donacobius – 1
               80. Family: Bernieridae – Malagasy Warblers – 11
               81. Family: Pnoepygidae – Cupwings – 4
               82. Family: Hirundinidae – Swallows – 88
            Superfamily: Sylvioidea  7/640
               83. Family: Pycnonotidae – Bulbuls – 156
               88. Family: Sylviidae – Sylviid Warblers & Allies – 32
               89. Family: Paradoxornithidae – Parrotbills – 38
               90. Family: Zosteropidae – White-eyes, Yuhinas & Allies – 148
               91. Family: Timaliidae – Tree-Babblers, Scimitar-Babblers & Allies – 58
               93. Family: Leiothrichidae – Laughingthrushes & Allies – 143
                     Family: Alcippeidae – Alcippe fulvettas  (now in Leiothrichidae)
               92. Family: Pellorneidae – Ground Babblers & Allies – 65
            Superfamily: Aegithaloidea – 4/128
               84. Family: Phylloscopidae – Leaf Warblers – 80
               85. Family: Hyliidae – Hylias – 2
               87. Family: Aegithalidae – Long-tailed Tits – 11
               86. Family: Scotocercidae – Bush Warblers & Allies – 35
                     Family: Cettidae – Cettia bush warblers & allies (now in Scotocercidae)  
                     Family: Erythrocercidae – Yellow flycatchers  (now in Scotocercidae)
         Parvorder Muscicapida – 19/856 (previously superfamily Muscicapoidea)
            Superfamily: Bombycilloidea – 6/15
               110. Family: Dulidae – Palmchat – 1
               107. Family: Bombycillidae – Waxwings – 3
               109. Family: Ptiliogonatidae – Silky-flycatchers – 4
               111. Family: Hylocitreidae – Hylocitrea – 1
               112. Family: Hypocoliidae – Hypocolius – 1
               108. Family: Mohoidae – Hawaiian Honeyeaters – 5
            Superfamily: Muscicapoidea – 8/693
               100. Family: Elachuridae – Spotted Elachura – 1
               101. Family: Cinclidae – Dippers – 5
               106. Family: Muscicapidae – Old World Flycatchers – 345
               105. Family: Turdidae – Thrushes & Allies – 175
               102. Family: Buphagidae – Oxpeckers – 2
               103. Family: Sturnidae – Starlings – 125
               104. Family: Mimidae – Mockingbirds & Thrashers – 34
               94. Family: Regulidae – Kinglets – 6
            Superfamily: Certhioidea  5/148
               95. Family: Tichodromidae – Wallcreeper – 1
               96. Family: Sittidae – Nuthatches – 29
               97. Family: Certhiidae – Treecreepers – 11
               98. Family: Polioptilidae – Gnatcatchers – 21
               99. Family: Troglodytidae – Wrens – 86
         Parvorder Passerida – 31/1721 (previously superfamily Passeroidea)
          Basal families in Passerida clade – 15/863
               113. Family: Promeropidae – Sugarbirds – 2
               114. Family: Modulatricidae – Dapple-throat & Allies – 3
               116. Family: Nectariniidae – Sunbirds & Spiderhunters – 148
               115. Family: Dicaeidae – Flowerpeckers – 53
               118. Family: Chloropseidae – Leafbirds – 12
               117. Family: Irenidae – Fairy-bluebirds – 3
               119. Family: Peucedramidae – Olive Warbler – 1
               120. Family: Urocynchramidae – Przevalski’s Pinktail – 1
               121. Family: Ploceidae – Weavers & Allies – 123
               123. Family: Viduidae – Indigobirds – 20
               122. Family: Estrildidae – Waxbills & Allies – 138
               124. Family: Prunellidae – Accentors – 12
               125. Family: Passeridae – Old World Sparrows – 43
               126. Family: Motacillidae – Wagtails & Pipits – 69
               127. Family: Fringillidae – Finches, Euphonias & Allies – 235
            Superfamily: Emberizoidea 16/858 (previously New World 9-primaried oscines)
               129. Family: Rhodinocichlidae – Thrush-Tanager – 1
               128. Family: Calcariidae – Longspurs & Snow Buntings – 6
               130. Family: Emberizidae – Old World Buntings – 44
               142. Family: Cardinalidae – Cardinals & Allies – 51
               141. Family: Mitrospingidae – Mitrospingid Tanagers – 4
               143. Family: Thraupidae – Tanagers & Allies – 384
               131. Family: Passerellidae – New World Sparrows – 132
               140. Family: Parulidae – New World Warblers – 115
               138. Family: Icteriidae – Yellow-breasted Chat – 1
               139. Family: Icteridae – Troupials & Allies – 106
               132. Family: Calyptophilidae – Chat-Tanagers – 2
               136. Family: Zeledoniidae – Wrenthrush – 1
               137. Family: Teretistridae – Cuban Warblers – 2
               134. Family: Nesospingidae – Puerto Rican Tanager – 1
               135. Family: Spindalidae – Spindalises – 4
               133. Family: Phaenicophilidae – Hispaniolan Tanagers – 4

The final extant species of the final family Phaenicophilidae (Hispaniolan Tanagers) in the Passerine list. 


Green-tailed Warbler (or Ground-Tanager) Microligea palustris.
Photo: John C Sullivan. eBird: Green-tailed Warbler 

That’s the end of it. Now all you have to do is go out and find all those Orders and Families and Genera and Species of birds.
I hope you found these ten postings as enlightening and entertaining as I did composing and presenting them. Don’t forget to check back in a year or so by which time everything will have changed utterly.

A few useful and informative links

https://avibase.bsc-eoc.org/familytree.jsp?lang=HA
https://checklist.americanornithology.org/taxa/
http://creagrus.home.montereybay.com/sylvid-new.html 
https://tree.opentreeoflife.org/opentree/opentree14.9@ott1041547/Passeriformes
https://en.wikipedia.org/wiki/Passerine
https://en.wikipedia.org/wiki/Passerine#Suborder_Acanthisitti
https://en.wikipedia.org/wiki/Passerine#Taxonomic_list_of_Passeriformes_families per IOU
https://en.wikipedia.org/wiki/Strisores 

http://www.j-avianres.com/en/article/doi/10.5122/cbirds.2013.0016
A review of the recent advances in the systematics of the avian superfamily Sylvioidea, Per ALSTRÖM Urban OLSSON, Fumin LEI

The Taxonomy Series

Installments post ever other day; installments will not open until posted.

Taxonomy One:  A brief survey of the history and wherefores of taxonomy: Aristotle, Linnaeus and his binomial system of nomenclature, taxonomic ranks and the discovery and application of biological clocks.
Taxonomy Two:  Introduces the higher levels of current taxonomy: the three Domains and the four Kingdoms. We briefly discuss Kingdom Protista, then the seven phyla of Kingdom Fungi.
Taxonomy Three:  Kingdom Plantae.
Taxonomy Four:  Kingdom Animalia to Phylum Annelida.
Taxonomy Five:  A discussion of Cladistics, how it works and why it is becoming ever more important.
Taxonomy Six:  Phylum Chordata, stopping at Class Mammalia.
Taxonomy Seven:  Class Mammalia.
Taxonomy Eight:  Class Aves, beginning with a comparison of five different avian checklists of the past 50 years.
Taxonomy Nine:  A cladogram and discussion of Subclass Neornithes (modern birds) of the past 110 million years, reaching down to the current forty-one orders of birds.
Taxonomy Ten:  A checklist of Neornithes including all ranks and clades down to the rank of the current 251 families of birds (plus a few probable new arrivals) with totals of the current 11,017 species of birds.

There will be a relatively brief look at “pecularities & problems of the checklist,” not yet written. Following that there may be some brief accounts of a few monotypic species which I think may be the most interesting birds.

Cladogram of forty-one Avian Orders | Taxonomy 9

September 10, 2024

[By Chuck Almdale]

Presented below is a cladogram of the forty-one currently recognized Orders of birds. A cladogram is a graphic representation of relationships between organisms with one of the axes, in this case the horizontal, capable of representing the passage of time, if the designer chooses to do so. For those unfamiliar with cladograms in general, Taxonomy 5 on cladistics will help; this cladogram’s significant features are also listed below. Frequent referral to the cladogram should clarify the descriptions of the clades.

The Cretaceous–Paleogene (K–Pg) extinction event


Asteroid strikes the earth 66 mya; not an actual photograph of the event. Wikipedia: K-Pg Extinction Event

Critically important in the evolution of birds was the Cretaceous–Paleogene (K–Pg) extinction event, perhaps better known by the prior term Cretaceous–Tertiary (K–T) extinction. (Cretaceous is spelled with a “K” in German). Approximately 66 million years ago an asteroid smashed into the earth where the Gulf of Mexico now lies, throwing an immense quantity of water, rock and dirt into the atmosphere. This material spread by winds around the earth, causing a “nuclear-winter”-type worldwide blackout that wiped out an estimated three-quarters of plant and animal species on earth.  All non-avian dinosaurs became extinct as did most tetrapods weighing over 55 pounds (25 kg), excepting a few cold-blooded species such as sea turtles and crocodilians. This marks the end of the Cretaceous period and the entire Mesozoic era and the beginning of the Paleogene period of the Cenozoic era, our current geological era. The asteroid impact left a thin layer of sediment – the K–Pg (K–T) boundary, found worldwide in marine and terrestrial rocks – which contains high levels of iridium, a metal more common in asteroids than in the Earth’s crust. Many of the avian clade evolutionary divergences displayed in the cladogram occurred at or very shortly after this event.

Features of the Cladogram

K-Pg (K-T) event: This is marked by a vertical broken orange line.

Timescale: At top and bottom, 110 million years ago on left, current day on right.

41 Orders: Listed on right side, generally the most ancient at top, the youngest at bottom, numbered on right margin. For three higher ranks the number of orders they contain is listed, e.g. Parvclass Neoaves – 34.

Families & Species: Listed next to the Order name, e.g. “Rheas  Rheiformes  1-2” means 1 family, 2 species within Order Rheiformes.

Heavy Dark Lines: The branches of the evolutionary tree. The tree starts near the upper left with Subclass Neornithes, appearing 110-101 million years ago, and branches right with branch tips ending at the current 41 Orders.

Nine Clades: In red. the same nine clades used in the chart of five checklists presented in Taxonomy 8.

The Diversification Cluster: Many major lineage divisions occurred within a few million years following the K-Pg extinction event.

The cladogram was created in MS Excel.

Overview of Class Aves to Subclass Neornithes

The biological Class Aves includes fossil groups of birds as well as modern taxa. It is generally considered to have emerged out of the Theropod (bipedal carnivorous) dinosaurs 150 million years ago (mya), with Archaeopteryx, the feathered, toothed and bony-tailed dinosaur as the first of its kind. Archaeopteryx could not lift its wings above its back, but could flap downward, and it’s now believed it could fly a bit like a pheasant, with short bursts of active flight as well as short glides. Recent X-ray studies revealed nearly hollow bones. It had asymmetrical feathers suggesting flight ability; flightless birds have symmetrical wing feathers (except for those lacking wings). Scientists are still debating how it got into the air: did it run, leap from a perch or wait for a strong breeze? The question of whether it should be considered a dinosaur or the first bird is also up in the air. Others avians appeared during the late Jurassic or early Cretaceous periods (150 – 110 mya); all are extinct and are placed in Subclass Archaeornithes (not on this cladogram). This leaves remaining the line leading to the as-yet-extant species, Subclass Neornithes, located at upper left in the cladogram below.

Subclass Neornithes includes all extant avian species. The earliest divergence within Neornithes, around 102 mya, split Infraclass Paleognathae (ratites and tinamous) from Infraclass Neognathae. Neognathae include two primary major groups of birds, Parvclass Galloanserae and Parvclass Neoaves. Galloanserae in turn is composed of two Orders, Anseriformes (waterfowl) and Galliformes (chickens and allies). All other modern birds fall into Parvclass Neoaves.

The information in the Subclass Neornithes cladogram below is relatively up-to-date (new information keeps pouring in), and comes from a variety of sources. The currently accepted 41 orders of bird appear roughly in sequence of oldest to youngest, and fall into nine major clades (in red). Credible dates of appearance  (e.g. 60 mya) of any clade can vary 5% (e.g. 63-57 mya) from dates indicated.

To print or save an image of the chart: click here.

Increase chart size: <Control> +; Decrease: <Control> -.

The rest of this post discusses the nine clades (above in red), each beginning with a excerpt from the main cladogram.

Subclass Neornithes – Infraclass Palaeognathae

About 102 million years ago, birds began to diversify into two major clades, the Palaeognathae and the Neognathae. The small clade of Palaeognathae began diversifying about 80 million years ago (mya) and is currently comprised of five of the most ancient orders of birds. The line leading to modern Struthioniformes (Ostriches) was the first to diverge 80 mya, followed by the Rheiformes (Rheas in South America) around 71 mya, then the Tinamiformes (Tinamous of Central and South America) at 69 mya, leaving Superorder Apterygimorphae, consisting of the Orders of Apterygiformes (Kiwis) and Casuariiformes (Cassowaries and Emu). These last two orders diverged from one another at the K-Pg extinction event 66 mya. The Emu later diverged (not shown here) from Cassowaries around 32 mya. Today the Palaeognathae contains 5 orders, 5 families and 59 species, 46 of them Tinamous. There are no monotypic families.


Somali Ostrich, getting closer to extinction.
Photo: Christiaan Kooyman, Jan. 2003. Wikipedia: Somali Ostrich

Cladogram used as source for Palaeognathae splits


Diagram from: Microstructural and crystallographic evolution of palaeognath (Aves) eggshells. Seung Choi, Mark E Hauber, Lucas J Legendre, Noe-Heon Kim, Yuong-Nam Lee, David J Varricchio. Jan 31, 2023.
Link: https://elifesciences.org/articles/81092 scroll to Fig. 13 about 1/3rd way down.

There is a potential problem with the Palaeognathae cladogram directly above, also depicted in the Palaeognathae portions of the Neornithes cladograms. Ostriches, Rheas, Cassowaries, Emu and Kiwis were long considered to be each other’s closest relatives and constituted the Ratites, flightless birds with a flat (not keeled) sternum (breastbone). The sternal keel – possessed by all flying birds – provides additional bone attachment surface for breast muscles. However, Tinamous are not flightless and possess a sternal keel: they don’t like to fly and prefer running or walking away silently into the forest or grassland, never to be seen but only heard, but can fly when necessary. But if the Tinamou line is embedded within the ratites – now described as “mostly flightless” in order to accommodate the capable-of-flight Tinamous – it means Tinamous developed their sternal keel separately from the Neognathae, the clade which includes all birds except the ratites and tinamous. The depicted schema also has Tinamous, Cassowaries, Emu and Kiwis diverging from Rheas around 70 mya. But Tinamous and Rheas are currently endemic to the Americas, while the other three groups live in Australia, New Guinea and New Zealand. Based on geography, one might assume that a Tinamou-Rhea group diverged from a Cassowary-Emu-Kiwi group, then each group continued their own divergences. Yet a 2010 study suggests that the extinct Moa of New Zealand is the Tinamou’s closest relative. All of these birds have a similar and distinctive Palaeognath palate. Supercontinent Gondwana didn’t finish breaking up into Africa, India, Australia, South America and Antarctica until after – perhaps well after – the start of the Paleogene (66 mya), so these flightless birds had many millions of years to walk to their current ranges from wherever in Gondwana they began. The current opinion holds that the ratites developed flightlessness several times; it may be that flight developed several times as well (Tinamous and Neognathae). However, if some of the dating studies are inaccurate, it may yet be that the Tinamous were first to diverge from the rest of the Palaeognaths (or vice-versa which amounts to the same thing, but suggests that the ratites developed from a flying ancestor), a possibility depicted occurring about 84 mya in the partial cladogram below.


Ordinal-level genome-scale family tree of modern birds. The Origin and Diversification of Birds. Stephen L. Brusatte, Jingmai K. O’Connor, and Erich D. Jarvis. Current Biology Review; Vol. 25, Issue 19; 5 Oct 2015, figure 6. [Link]

What these Palaeognaths do have in common is indicated by their name which means “ancient jaw”: the bones of their palate are described as retaining basal (primitive) morphological characteristics, thus closer to the reptilian palate than that of the birds in Infraclass Neognathae (“new jaw”).

It seems there may still be details to be worked out with the Palaeognathae relationships among themselves and with the Neognathae. “Early days” perhaps, as they say in BBC police shows, meaning “We’re still working on it, please stop asking and go away.”

Subclass Neornithes – Infraclass Neognathae

The clade Infraclass Neognathae is the sister taxon to Infraclass Palaeognathae. This clade contains the remaining 36 orders of birds and the rest of the approximately 11,000 (a perpetually increasing number) species of birds. This clade – some say Infraclass, others say Parvclass and still others simply leave it as a clade – diverged from the Palaeognathae 102 mya and began diversifying into two clades around 88 million years ago. The smaller of these two clades is Parvclass Galloanserae, consisting of two orders and 483 species, and the much larger Parvclass Neoaves consisting of the remaining 34 orders.

Subclass Neornithes – Infraclass Neognathae – Parvclass Galloanseres

Parvclass Galloanserae diverged from the rest of Neognathae 88 million years ago, when a major breakup within the supercontinent Gondwana occurred. The Galloanseres lineage began their diverging into two orders at the start of the Paleogene Period (66-23 mya): Anseriformes  consisting of Waterfowl, Screamers (South America) and the Magpie-Goose (Australia), 3 families totaling 178 species; the other is Galliformes, a diverse worldwide order containing Pheasants, Quail, Guineafowl (Africa), Guans (Central & South America) and Megapodes (Australasia), 5 families  totaling 305 species. The Megapodes have the unique behavior of not sitting on their eggs but using warm earth, fermenting vegetation or geothermal heat to incubate them. Later Galloanserae diversification followed their dispersal to the various continents and islands during the Eocene (56-33.9 mya).  The monophyletic (one lineage) relationship of these two orders and their placement as the closest relative (sister taxon) of Neoaves are well supported. Background: Waterfowl and Gamefowl (Galloanserea); Pereira, Sergio L. & Baker, Allan J.; page 416, Link: Timetree.temple

The largest families are Phasianidae (Pheasants & allies, 186 species) and Anatidae  (Ducks & allies, 174 species). Family  Anseranatidae (Magpie Goose of Australia) is monotypic.

Magpie Goose (Anseranas semipalmata), a monotypic family of northern Australia and southern New Guinea.
Photo: JJ Harrison, Dec. 2019. Wikipedia: Magpie Goose

As mentioned previously, molecular clocks aren’t the only factor in drawing up cladograms which include dates on the nodes of divergence; morphology is still important, especially when it comes to calibrating a lineage’s molecular clock. Following are two examples of how morphological studies are still being used.

Vegavis the Neoornithine


Vegavis fossil from Letters from Gondwana. Source: Paleonerdish  OK

Although a Cenozoic origin for Galloanserae has been long hypothesized based on fragmentary and incomplete specimens, definitive evidence was found only recently. Vegavis iaii, the oldest known anseriform fossil from the Maastrichtian stage (72.1-66 mya) of the late Cretaceous, is closely related to the lineage of ducks and geese. This finding implies that modern anseriform families, and hence their closest living relative, the Galliformes, were already independent lineages in the late Cretaceous. From: Waterfowl and Gamefowl (Galloanserea); Pereira, Sergio L. & Baker, Allan J.; page 416, Link: Timetree.temple 

The single best record of a Cretaceous neornithine is the partial skeleton of Vegavis from the last sliver of the  Cretaceous period (around 68–66 million years ago) in Antarctica. This bird is assigned to the Neornithes  subgroup of modern birds known as Galloanserae which includes Anseriformes  (ducks and geese) based on the morphology of its well-developed hypotarsus.
Source: The Origin and Diversification of Birds. Stephen L. Brusatte, Jingmai K. O’Connor, and Erich D. Jarvis. Current Biology Review; Vol. 25, Issue 19; 5 Oct 2015, page. [Link]

The Hypotarsus


Hypotarsus process on the tarsometatarsus. Pinterest: Mark Kobelka 

The hypotarsus is a process [bony bump] on the posterior side of the upper end of the tarsometatarsus (the long bone immediately above the foot) of many birds; aka the calcaneal process. A protruding hypotarsus “enhances the capacity of diving birds to propel their feet backward and power underwater swimming.
Source: Research Gate, Comparative hindlimb myology of foot‐propelled swimming birds, article by Glenna T. Clifton, Jennifer A Carr & Andrew A Biewener.   

The Avian Quadrate Bone

We previously discussed the avian quadrate bone in the first and eighth posting in this series, but it doesn’t hurt to be reminded of this major marker in the early evolution of birds. You’ll never look at ducks and chickens the same way again.


Mallard skull and quadrate anatomy. (A) Lateral view, (B) Ventral view. Bones: Quadrate (red), Pterygoid (purple), Palatine (green), Jugal (blue), Upper Bill. From: Kinematics of the Quadrate Bone During Feeding in Mallard Ducks. [Link] (2011) Megan M Dawson, Keith A Metzger, David B Baier & Elizabeth L Brainerd.  [Diagram used in Taxonomy 1 & 8.]

In birds and [other] reptiles, the quadrate acts as a hinge between the lower jaw and the skull and plays an important role in avian cranial movement. In a study by Kuo, Benson and Field (2023), the quadrate of 50 currently living galloanseran species were examined resulted in a reconstruction of the ancestral Galloanserae which closely approximated the average of living modern galloanserans. But when they added early fossil galloanseran quadrates to the study the results shifted, indicating that the quadrate of the common ancestor of galliforms and anserforms was closer to the quadrate of living galliforms than that of living anserforms. In short, the common ancestor of all chickens and ducks was [likely] more like a chicken than a duck. Morphology still has important things to add.

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves

The remaining 34 orders and the rest of almost 10,500 species of birds are in the Parvclass clade of Neoaves, which we will discuss as seven clades. All of these clades appeared 60-70 mya, close to the end of the Cretaceous era and start of the Paleocene era 66 mya, caused by that cataclysmic impact of an asteroid into what is now the Gulf of Mexico, north of the Yucatan peninsula. Except for the warm-blooded birds, the dinosaurs vanished and the Neoavian evolutionary radiation exploded. All dates are molecular clock calculations, based not on proteins but on ~30 million pairs of genomic (chromosomal) DNA. There are many competing systems for Neoaves phylogeny, this site shows ten of them.

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Columbea

The Clade Columbea currently consists of 5 orders, 5 families and 400 species. Their first divergence was the widespread Phoenicopterimorphae Superorder (orders of flamingos and grebes) from Columbimorphae at 68 mya. Within Columbimorphae Order Columbiformes (Pigeons & Doves) diverged at 65 mya, followed at 62 mya by the divergence between Order Mesitornithiformes (Mesites of Madagascar) and Order Pterocliformes (Sandgrouse, ranging from east Asia to southern Africa). [Some scientists do not combine Phoenicopterimorphae  and Columbimorphae into a single clade; one such alternative suggest two clades, Mirandornithes (Flamingos & Grebes) and Columbimorphae (Pigeons, Mesites and Sandgrouse) as two simultaneously emerging clades, with sister taxon Passerea containing all remaining birds. This creates a polytomy which in the current view cannot be correct.] Several of the following cladesSome scientists do not combine Phoenicopterimorphae  and Columbimorphae into a single clade. Several of the following clades are larger with more divergences. If you’ve compared the text so far with the cladogram(s), you should have a feeling for the grouping and divergences and won’t need all the additional detailed descriptions as have been supplied so far. The largest family in Clade Columbea by far is Columbidae (Pigeons & Doves, 353 species). The smallest family is Mesitornithidae (Mesites of Madagascar, 3 species).

Subdesert Mesite (Monias benschi) of southwestern Madagascar.
Photo: Ben Rackstraw, Nov. 2006. Wikipedia: Mesite

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea

After the divergence of the clade Columbea, the remaining Neoaves are tentatively classified into the clade Passerea. This encompasses all the remaining 29 orders, 233 families and 10,075 species.  We will discuss these as six clades.

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea Clade Otidae

The first group within this large clade of Passerea consist of the Clade Otidae. This contains superorders Caprimulgimorphae & Otidimorphae, which combined comprise 4 orders, 11 families and 805 species. 609 of these species are in the Order Caprimulgiformes, an increasingly diverse order comprising 8 families: Hummingbirds, Treeswifts, Swifts, Owlet-Nightjars, Oilbird, Potoos, Nightjars and Frogmouths. Fifty years ago there were only 105 species in 5 families classified to Caprimulgiformes. These two superorders diverged from the rest of Clade Passerea 68 mya, the same time that Superorder Phoenicopterimorphae diverged from the rest of Clade Columbea (described above). With the 5% (+/- 3.4 million years) credible dates (margin of error), this was when the asteroid extinction even occurred. During the very early Paleogene (65-61 mya) this clade diverged into its current four Orders of Caprimulgiformes (Hummingbirds, Swifts, Nightjars, Potoos & allies),  Cuculiformes (Cuckoos), Musophagiformes (Turacos of Sub-Saharan Africa) and Otidiformes  (Bustards – widespread but uncommon in the Old World). The largest families are Trochilidae (Hummingbirds, 155 species in the Americas), Cuculidae (Cuckoos, 147 species worldwide) and Caprimulgidae (Nightjars and allies, 97 species worldwide). The only monotypic family is: Steatornithidae (Oilbird of the American tropics).


Oilbirds (Steatornis caripensis) nest and roost in caves across northern and northwestern South America.
Photo: The Lilac Breasted Roller. April 2007. Wikipedia: Oilbird

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea – Clade Gruae

The clade Gruae appeared at the K-Pg extinction event, 66 mya. Within two million years it had already begun diversifying into two Superorders, Opisthocomimorphae and Gruimorphae. Opisthocomimorphae today consists of a single family Opisthocomidae with a single species (the Hoatzin of South America). Gruimorphae diverged into two orders: Gruiformes (six families of  Flufftails, Coots, Finfoots, Limpkin, Trumpeters and Cranes – 192 species worldwide), and the very diverse Charadriiformes  (19 families of Gulls, Sandpipers, Plovers, Alcids, Buttonquail, Pratincoles, Oystercatchers, Thick-knees and 11 other families of under ten species each – 386 species worldwide). The largest families are Rallidae (Rails, 155 species), Laridae (Gulls and allies, 100 species) and Scolopacidae (Sandpipers, 97 species). There are six monotypic families: Opisthocomidae (Hoatzin of South America), Aramidae (Limpkin So.-Cent. America), Pluvianellidae (Magellanic Plover of Southern South America), Ibidorhynchidae (Ibisbill of South Asian mountains), Pedionomidae (Plains-wanderer of eastern Australia) and Dromadidae (Crab-Plover around the Indian Ocean).


Ibisbill (Ibidorhyncha struthersii) , a bird of central Asian mountain streams.
Photo: Mohanram Kemparaju, March 2008. Wikipedia: Ibisbill

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea – Clade Eurypygimorphae (or Phaethontimorphae)

Clade Eurypygimorphae is a small clade consisting of the two Orders Eurypygiformes and Phaethontiformes, two families and only 5 extant species. It is the sister taxon to Clade Aequornithes as their common ancestor appeared shortly after the K-Pg extinction event, and these two clades began to separate within a million years after that. The reason I denote it as a separate clade is because an important source report (Brusette, O’Conner, Jarvis 2015 The origin and diversification of birds; link) classified them as within Clade Passerea, but not within the clade of Core Waterbirds (Aequornithia), thus indicating that they were a basal clade. The two orders within this clade diverged from each other 5 million years later, or 60 mya. Order Eurypygiformes contains only the Kagu of New Caledonia and the Sunbittern of the American tropics, two widely-separated locations. Order Phaethontiformes contains three species of Tropicbirds, previously classified to the order Procellariformes next to the Pelicans.

Sunbittern (Eurypyga helias) of Central and northern South America, displaying its “suns.” Stavenn, Jan 2007. Wikipedia: Sunbittern

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea – Clade  Aequornithes (Core Waterbirds)

Clade Aequornithes (or Aequornithia) or Core Waterbirds consists of three sub-clades, six orders, 16 families and 365 species. Clades Aequornithes and sister taxon  Eurypygimorphae diverged from the final two clades of Afroaves and Australaves, see below) very soon after the K-Pg extinction event. Aequornithes separated from Eurypygimorphae about 1 million years later. The loons then diverged, soon followed by divergence of the Stork, Cormorant and Pelican Orders from the Penguins and Tubenoses, 61 mya. Within a few million years these clades had further diversified into the six current orders. The largest families are Procellariidae (Shearwaters and Petrels – 98 species worldwide) and Ardeidae (Herons, Egrets, and Bitterns – 71 species worldwide). There are two monotypic families: Balaenicipitidae (Shoebill of Central Africa), and Scopidae (Hamerkop of sub-Saharan Africa).

Hamerkop (Scopus umbretta) of sub-Saharan Africa, southwest Arabia and Madagascar. This 1 lb. 22 inch tall bird builds one of the largest nests in the world; one was estimated to contain 8,000 sticks.
Photo: Sumeet Moghe, August 2012. Wikipedia: Hamerkop

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea – Clade Telluraves (Core Landbirds)

Clade Telluraves is the sister taxon to Clade Aequornithes and has two main branches, the sister taxa of Clade Afroaves and Clade Australaves. These two clades diverged 63 mya.

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea – Clade Telluraves (Core Landbirds) – Clade Afroaves

Our last two clades and sister taxa of Afroaves and Australaves diverged from one another 63 mya (+/- 3.2 my). Looking at the cladogram you’ll see that Clade Afroaves diverged up to seven more times before reaching the rank of Order. The clade’s current ten orders evolved 63-42 mya and now totals 28 families (11.2% of 251 families) and 1,254 species (11.4% of 11,017 species). [Those two percentages seem oddly similar.] The three largest families by far are: Accipitridae (Hawks, Eagles & Kites – 250 species worldwide), Picidae (Woodpeckers – 235 species worldwide), and Strigidae (Owls – 229 species worldwide). Three families are monotypic: Pandionidae (everyone’s favorite land-based fish-eater, the Osprey, found worldwide), Sagittariidae (the Secretarybird of sub-Saharan Africa) and Leptosomidae (the Cuckoo-roller, a Madagascar endemic). Cuckoo-Roller is the sole denizen of one of the two monotypic orders; the other is the Hoatzin of Opisthocoformes.

Cuckoo-Roller (Leptosomus discolor) of Madagascar an Comoros Islands, female or juvenile; named for its resemblance to both cuckoos and rollers.
Photo: frank wouters, October 2005. Wikipedia: Cuckoo-Roller

Subclass Neornithes – Infraclass Neognathae – Parvclass Neoaves – Clade Passerea – Clade Telluraves (Core Landbirds) – Clade Australaves

Our last clade, in some ways the most unusual, diverged from its sister clade Afroaves 63 mya (+/- 3.2 my), shortly after that infamous K-Pg asteroid strike. The small Order of Cariamiformes (Seriemas, 2 species endemic to the grasslands of central South America) diverged perhaps less than a million years later from, leaving Clade (or Superorder) Eufalconimorphae (falcons, parrots & passerines) remaining. Falconiformes  (Falcons & Caracaras, 65 species worldwide, all in one family) diverged 60 mya, leaving clade Psittacopasseres, a portmanteau name encompassing the two remaining Orders of Psittaciformes and Passeriformes. As Cariamiformes and Falconiformes are more basal (closer to the root) than Psittacopasseres, it suggests that the last common ancestor of the entire clade had a predatory lifestyle. The final divergence 55 mya was between sister taxa Psittaciformes (Parrots & Cockatoos, 405 species in 4 families, worldwide) and Passeriformes. Passeriformes (songbirds) is an enormous order: a whopping 143 families [57% of 251 total families] and a stunning 6,595 species [60% of 11,017 species], 49% more species than all the 40 other orders combined). It is again interesting to note that the percentages for family and species diversity are nearly the same. The size of this order – 60% of all bird species – explains why most birders frequently speak of birds as divided into only two groups: passerines and non-passerines.

Blue-banded Pitta (Erythropitta arquata) of Borneo, a beautiful member of a beautiful and reclusive family.
Photo: JJ Harrison. Wikipedia: Blue-banded Pitta 

Our next and penultimate posting in this series will focus on the huge and hugely diverse Order Passeriformes.

The Taxonomy Series

Installments post ever other day; installments will not open until posted.

Taxonomy One:  A brief survey of the history and wherefores of taxonomy: Aristotle, Linnaeus and his binomial system of nomenclature, taxonomic ranks and the discovery and application of biological clocks.
Taxonomy Two:  Introduces the higher levels of current taxonomy: the three Domains and the four Kingdoms. We briefly discuss Kingdom Protista, then the seven phyla of Kingdom Fungi.
Taxonomy Three:  Kingdom Plantae.
Taxonomy Four:  Kingdom Animalia to Phylum Annelida.
Taxonomy Five:  A discussion of Cladistics, how it works and why it is becoming ever more important.
Taxonomy Six:  Phylum Chordata, stopping at Class Mammalia.
Taxonomy Seven:  Class Mammalia.
Taxonomy Eight:  Class Aves, beginning with a comparison of five different avian checklists of the past 50 years.
Taxonomy Nine:  A cladogram and discussion of Subclass Neornithes (modern birds) of the past 110 million years, reaching down to the current forty-one orders of birds.
Taxonomy Ten:  A checklist of Neornithes including all ranks and clades down to the rank of the current 251 families of birds (plus a few probable new arrivals) with totals of the current 11,017 species of birds.

A FEW USEFUL & INFORMATIVE SITES:

https://en.wikipedia.org/wiki/Aequornithes 
https://en.wikipedia.org/wiki/Afroaves 
https://en.wikipedia.org/wiki/Australaves 
https://en.wikipedia.org/wiki/Austrodyptornithes  
https://en.wikipedia.org/wiki/Cavitaves 
https://en.wikipedia.org/wiki/Eufalconimorphae
https://en.wikipedia.org/wiki/Gruae 
https://en.wikipedia.org/wiki/Otidimorphae 
https://en.wikipedia.org/wiki/Passerea
https://en.wikipedia.org/wiki/Pelecanimorphae 
https://en.wikipedia.org/wiki/Phaethoquornithes
https://en.wikipedia.org/wiki/Picocoraciae 
https://en.wikipedia.org/wiki/Strisores
https://en.wikipedia.org/wiki/Telluraves 

Volunteers needed at Audubon Ballona Wetlands Education Program

September 9, 2024

[Posted by Chuck Almdale, submitted by Cindy Hardin]

Nature Nexus Institute has taken over the operation of Los Angeles Audubon’s school outreach program of school field trips to the Ballona Wetlands. Santa Monica Bay Audubon Society has helped support this organization for several decades.

Nature Nexus Institute is all about sharing and teaching local school children about the special habitats found right here in Los Angeles. They will be starting their six-week Fall Training on 17 September to get ready for school field trips to the Ballona Wetlands. Their tours take place on Tuesdays and Thursdays. All you need to be a volunteer is a love of the outdoors and the enthusiasm to work with school age aspiring nature lovers. Please contact Cindy Hardin at chardin@naturenexusinstitute.org or give her a call at 310-745-2118 if you are interested.

The following announcement is from them.

One of the Ballona Salt Marsh channels (Leslie Davidson ’07)

We are looking for Volunteers!

I have begun the process of booking field trips, which means Fall Training is on the horizon. We have a great group of speakers, some of whom are brand new to the program, and I am excited to hear what they have to share about all things Nature, History and Ballona.
 
Tuesday dates: Tuesdays 9/17, 9/24, 10/1, 10/8, 10/15, 10/22
Time:  9 am to noon, unless otherwise noted (see below)
Location: Ballona Wetlands Ecological Reserve, entrance at 303 Culver Blvd., Playa del Rey.
Contact: Cindy Hardin, Ph/Text 310-745-2118, <
chardin@naturenexusinstitute.org
>
 
The training schedule is as follows:

September 17thWelcome back and welcome newcomers, an overview of the program, the mission of Nature Nexus institute, description of Learning Stations and route of field trips. Speaker: me, Cindy Hardin!

September 24thWetland Ecology by speaker Dr. Dave Bader. Dave handles education at the Marine Mammal Care Center in San Pedro, and has lots of knowledge to share about the unique and vanishing wetland habitats of our coast. He is new to our roster, and I hear he is an excellent speaker!

October 1stProgram history and goals, Restoration Ecology and Education by Dr. Margot Griswold. Margot has been the driving force behind our work at both the Baldwin Hills and Ballona, and has done numerous restoration projects throughout the region. Her knowledge of native plants and habitats is unparalleled! She has also done huge advocacy work to spread knowledge to underserved communities-our target audience!

October 8thTo be determined. I am hoping that Greg Pauly, curator of Herpetology at the Natural History Museum will be available, but I am waiting to hear back from him. I will keep you all posted. If Greg is not available we will fill in the slot with another program. If anyone has a suggestion, I am all ears!

October 15thGabrieleno History, Practices and Culture by Matthew Teutimez. Mr. Teutimez has a wealth of knowledge about local indigenous culture, and he is a very compelling speaker. I have seen a couple of his talks, and they are quite informative and memorable. He will be very helpful in showing us what we can best communicate to our student visitors about local Native Peoples. PLEASE NOTE: OUR START TIME FOR THIS SESSION WILL COMMENCE ONE HOUR LATER THAN USUAL, AT 10 AM.

October 22nd – Birds and Birding with our own Walter Lamb. Walter is a fantastic birder, and patient and generous in sharing his skills and knowledge. He has a huge following for his Bird Walks that he leads during our Open Wetlands events, and I am so glad he has the time to work with our group.

Thanks,
Cindy